I am experiencing identical symptoms. It's as if there's a "snap" turned on to both the X and the Y axis.
I've confirmed that this does not exist at the hardware or driver level. Mouse pointer moves fluidly and smoothly across X and Y axis UNTIL THE ROM IS RUN VIA MAME.
It's very easy to see what's happening, because MAME turns on the crosshairs in any lightgun game.
I noticed this in Operation Wolf and also in Zero Point before I (1) tried unsuccessfully to find the setting to fix it, and (2) did a google search and landed here.
It can't be pointer specific at this point, but for what it's worth I am using a Wii controller with IR input turned on, using the CWiiD bundle for a pointer driver.
Again, the pointer works 100% perfectly in all applications except mame. :-(
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